Bed in Ref. 27 and aligned using a referencefree method (28) to generate the class averages shown in Fig. 6C. Using the in-plane orientations from the untilted particles, a threedimensional electron density map, with an estimated resolution of 29 ? was generated for both classes using the tilted particles, and these are shown in Fig. 7. As indicated by the original images, these maps show ring-sha
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